What are dung beetles?
Well, beetles that feed on dung. But while this is largely true, the definition of dung beetle is actually more complicated, being a mixture of an ecological trait (coprophagy) and a taxonomic classification (belonging to the scarabs). So dung beetles are those species of the large superfamily of scarab beetles (Scarabaeoidea) that feed on manure or belong to a taxonomic group that contains mainly dung-feeders. Dung feeding (coprophagy) has evolved several times within the scarabs, with the major groups being the Geotrupidae (dor beetles; ~150 species), the Aphodiinae (dung dwellers; ~3,500 species, Figure 1), and the Scarabaeinae (the dung beetles proper; ~7,000 species, Figure 2).
